A Revision of Pachacutia, a New Genus of Rare Andean Ithomiine Butterﬂies (Nymphalidae : Ithomiinae), with the Description of Two New Species
Willmott, Keith R.
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A newgenus, Pachacutian. gen., of large, rare Andean ithomiine butterßies is described and characterized, containing four species. Two species and their constituent subspecies and synonyms (Ithomia cleomella Hewitson, Ithomia mantura Hewitson, Ithomia honrathi Srnka, and Dircenna barrettii Dannatt) are transferred to Pachacutia from Godyris Boisduval (n. comb.). Two additional species are described from southern Ecuador (Pachacutia baroni n. sp.) and Bolivia (Pachacutia germaini n. sp.), and a new subspecies, Pachacutia mantura joroni n. ssp., is described from Peru. Pachacutia is a member of a clade containing otherwise small, often abundant lowland forest ithomiines, includingMcclungiaR.M. Fox, Brevioleria Lamas, and some species of Hypoleria Godman & Salvin. A comparative phylogenetic análisis showed Pachacutia species to be related as follows: (baroni (mantura (cleomella germaini))). A key for identiÞcation of all taxa, and illustrations ofmale and female genitalia for all species and adult specimens of all taxa, are presented, and the taxonomy and natural history of all taxa are discussed.
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